Cytoplasmic dynein has two heavy chains with globular heads that walk along the microtubule, to which they are bound by the stalks. Dynactin (not shown) may help attach the light chains to the cargo. Interactions between the stalks and the microtubule must repeatedly form and break.

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Cytoplasmic dynein has two heavy chains with globular heads that walk along the microtubule, to which they are bound by the stalks. Dynactin (not shown) may help attach the light chains to the cargo. Interactions between the stalks and the microtubule must repeatedly form and break.